Phylogenetic Relationships within Heliodinidae and Systematics of Moths Formerly Assigned to Heliodines Stainton (Lepidoptera: Yponomeutoidea): Yponomeutoidea)

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University of California Press, Jan 1, 2005 - Science - 205 pages
Heliodinids are tiny, brightly colored dayflying moths. Phylogenetic relationships among genera of Heliodinidae are proposed using parsimony and character compatibility. We describe and illustrate 45 North and Central species (25 newly named) assigned to five genera (two new, two exhumed from synonymy). Larval host plants are recorded for 33 species (14 newly discovered), about 45% of the known fauna; 90% of these are specialists on Caryophyllales, primarliy Nyctaginaceae.
 

Contents

Heliogemma preclara Hsu new species Figs 16 131200
79
PSEUDASTASIA Walsingham Figs 34 87 197
80
KEY TO SPECIES OF Euheliodines
81
Euheliodines jaliscella Hsu new species Fig 17 89 202
83
AETOLE Chambers revised status
84
Eximia Group Figs 18 3649 9091 133 203204
85
Aetole eximia Hsu new species Figs 18 3649 90 133 203
86
Aetole insolita Hsu new species Figs 1891204
88

KEY TO THE GENERA OF HELIODINIDAE AND SPECIES GROUPS OF AETOLE
31
Heliodines assemblage
33
Heliodines roesella Linnaeus Figs 72659 108 173
34
EPICROESA Meyrick Figs 27 60 109
37
Lamprolophus assemblage
38
Heliodines princeps Meyrick Figs 63 112 175
39
Lithariapteryx subassemblage
40
KEY TO SPECIES OF Neoheliodines
42
Neoheliodines eurypterus Hsu new species Figs 8 66 115 178
45
Neoheliodines megostiellus Hsu new species Figs 968 117 180
48
Neoheliodines melanobasilarus Hsu new species Figs 969 118 181
49
Neoheliodines vernius Hsu new species Figs 10 70 119 152154 157 163 182
50
Neoheliodines cliffordi Harrison and Passoa new combination Figs 1171 120183
53
Neoheliodines arizonense Hsu new species Figs 1172121 184
55
Neoheliodines albidentus Hsu new species Figs 1073 185
57
Aetole subassemblage
58
KEY TO SPECIES OF Embola OF NORTH AND CENTRAL AMERICA
60
Embola ionis Clarke new combination Figs 12 32 74 122 186
61
Embola ciccella Barnes and Busck new combination Figs 1375 123 161162 187
63
Embola cyanozostera Hsu new species Figs 144476 124 188
65
Embola albaciliella Busck new combination Figs 1477 189
66
Embolafriedlanderi Hsu new species Figs 1578125190
68
Embola melanotela Hsu new species Figs 1580127191
69
Embola autumnalis Hsu new species Fig 1581192
70
Embola sexpunctella Walsingham new combination Figs 82 193
71
Embola dentifer Walsingham Figs 13 83194
72
Embola powelli Hsu Figs 14 84 128 167 170 195
73
Embola xanthocephala Walsingham Figs 13 85 196
75
HELIOGEMMA Hsu new genus Figs 16 33 86 129131 8789
76
Heliogemma gigantea Hsu new species Figs 16 33 86 129 198
77
Heliogemma grandis Hsu new species Figs 16130 199
78
Bella Group Figs 1921 374748 9295 134137 205208
89
Aetole bella Chambers revised combination Figs 19374792134205
90
Aetole schulzella Fabricius new combination Figs 2093 135206
95
Aetole fulgida Hsu new species Figs 20 94 136 165 171 207
97
Aetole prenticei Hsu new species Figs 214895 137159208
98
Tripunctella Group Figs 2223 38 5154 96100 138142 209214
101
Aetole tripunctella Walsingham new combination Figs 22 385196 138209
103
Aetole unipunctella Walsingham new combination Figs 235297 139 172210
105
Aetole inusitata Hsu new species Figs 140211
109
Aetole aprica Hsu new species Figs 98 212
110
Aetole favonia Hsu new species Figs 2254 100 142214
112
Extraneella Group Figs 24 39 50 101 143 150 152 215
113
Aetole extraneella Walsingham new combination Figs 24 39 50 101 143 150 152 215
114
Calcifer Group Figs 2540 55 102103 144145 216217
118
Aetole calcifer Walsingham new combination Figs 25 40 55 102 144 216
119
Aetole calciferoides Hsu new species Figs 25 103 145 217
121
Galapagoensis Group Figs 4156 104 146218
123
Aetole galapagoensis Heppner and B Landry New combination Figs 4156 104 146 218
124
COPOCENTRA Meyrick Figs 43 105 148219
125
SCELORTHUS Busck Figs 4246 106 147 155 160 166 169 220
126
Heliodines marginata Walsingham Fig 107
127
Heliodines urichi Busck Fig 149
128
BIOLOGY
129
I
131
II
132
III
133
SUMMARY
135
IV
136
LITERATURE CITED
150
Figures 26220
161
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About the author (2005)

Yu-Feng Hsu is Associate Professor of Life Sciences at National Taiwan Normal University. Jerry A. Powell is Professor Emeritus of Entomology at the University of California, Berkeley and Director Emeritus of the Essig Museum of Entomology.

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